The domain within your query sequence starts at position 4309 and ends at position 4376; the E-value for the 1iur domain shown below is 5e-39.

ILKEVTSVVEQAWKLPESERKKIIRRLYLKWHPDKNPENHDIANEVFKHLQNEINRLEKQ
AFLDQNAD

This is a PDB domain - please see the PDBSUM entry of 1iur for further annotation